diff --git a/Lib/fontTools/feaLib/ast.py b/Lib/fontTools/feaLib/ast.py
index b0f805a7e..f998e306c 100644
--- a/Lib/fontTools/feaLib/ast.py
+++ b/Lib/fontTools/feaLib/ast.py
@@ -109,7 +109,7 @@ def asFea(g):
     if hasattr(g, 'asFea'):
         return g.asFea()
     elif isinstance(g, tuple) and len(g) == 2:
-        return asFea(g[0]) + "-" + asFea(g[1])   # a range
+        return asFea(g[0]) + " - " + asFea(g[1])   # a range
     elif g.lower() in fea_keywords:
         return "\\" + g
     else:
diff --git a/Tests/feaLib/data/ChainPosSubtable.fea b/Tests/feaLib/data/ChainPosSubtable.fea
index 46506229a..e64f88678 100644
--- a/Tests/feaLib/data/ChainPosSubtable.fea
+++ b/Tests/feaLib/data/ChainPosSubtable.fea
@@ -1,7 +1,7 @@
 feature test {
-    pos X [A-B]' -40 B' -40 A' -40 Y;
+    pos X [A - B]' -40 B' -40 A' -40 Y;
     subtable;
     pos X A' -111 Y;
     subtable;
-    pos X B' -40 A' -111 [A-C]' -40 Y;
+    pos X B' -40 A' -111 [A - C]' -40 Y;
 } test;
diff --git a/Tests/feaLib/data/GSUB_6.fea b/Tests/feaLib/data/GSUB_6.fea
index 82fdac250..223067016 100644
--- a/Tests/feaLib/data/GSUB_6.fea
+++ b/Tests/feaLib/data/GSUB_6.fea
@@ -1,10 +1,10 @@
 lookup ChainedSingleSubst {
     sub [one two] three A' by A.sc;
-    sub [B-D]' seven [eight nine] by [B.sc-D.sc];
+    sub [B - D]' seven [eight nine] by [B.sc - D.sc];
 } ChainedSingleSubst;
 
 lookup ChainedMultipleSubst {
-    sub [A-C a-c] [D d] E c_t' V [W w] [X-Z x-z] by c t;
+    sub [A - C a - c] [D d] E c_t' V [W w] [X - Z x - z] by c t;
 } ChainedMultipleSubst;
 
 lookup ChainedAlternateSubst {
diff --git a/Tests/feaLib/data/GSUB_8.fea b/Tests/feaLib/data/GSUB_8.fea
index 62c7bee4c..8f417496f 100644
--- a/Tests/feaLib/data/GSUB_8.fea
+++ b/Tests/feaLib/data/GSUB_8.fea
@@ -2,7 +2,7 @@ languagesystem DFLT dflt;
 
 feature test {
     rsub [a A] [b B] [c C] q' [d D] [e E] [f F] by Q;
-    rsub [a A] [b B] [c C] [s-z]' [d D] [e E] [f F] by [S-Z];
+    rsub [a A] [b B] [c C] [s - z]' [d D] [e E] [f F] by [S - Z];
 
     # Having no context for a reverse chaining substitution rule
     # is a little degenerate (we define a chain without linking it
diff --git a/Tests/feaLib/data/bug514.fea b/Tests/feaLib/data/bug514.fea
index 26da86557..1ef5af6ae 100644
--- a/Tests/feaLib/data/bug514.fea
+++ b/Tests/feaLib/data/bug514.fea
@@ -5,7 +5,7 @@
 # makeotf produces {A:-40, B:-40, C:-40} and {A:-111, B:-40} which
 # is redundant. https://github.com/adobe-type-tools/afdko/issues/169
 feature test {
-    pos X [A-B]' -40 B' -40 A' -40 Y;
+    pos X [A - B]' -40 B' -40 A' -40 Y;
     pos X A' -111 Y;
-    pos X B' -40 A' -111 [A-C]' -40 Y;
+    pos X B' -40 A' -111 [A - C]' -40 Y;
 } test;
diff --git a/Tests/feaLib/data/language_required.fea b/Tests/feaLib/data/language_required.fea
index 4005a781b..687c48a1c 100644
--- a/Tests/feaLib/data/language_required.fea
+++ b/Tests/feaLib/data/language_required.fea
@@ -18,5 +18,5 @@ feature liga {
 } liga;
 
 feature scmp {
-  sub [a-z] by [A.sc-Z.sc];
+  sub [a - z] by [A.sc - Z.sc];
 } scmp;
diff --git a/Tests/feaLib/data/spec4h1.fea b/Tests/feaLib/data/spec4h1.fea
index b43e13b0d..a3d249468 100644
--- a/Tests/feaLib/data/spec4h1.fea
+++ b/Tests/feaLib/data/spec4h1.fea
@@ -8,7 +8,7 @@ languagesystem latn TRK;
 languagesystem cyrl dflt;
 
 feature smcp {
-    sub [a-z] by [A.sc-Z.sc];
+    sub [a - z] by [A.sc - Z.sc];
 
     # Since all the rules in this feature are of the same type, they
     # will be grouped in a single lookup.  Since no script or language
diff --git a/Tests/feaLib/data/spec5f_ii_2.fea b/Tests/feaLib/data/spec5f_ii_2.fea
index b20a74c3b..916f797a8 100644
--- a/Tests/feaLib/data/spec5f_ii_2.fea
+++ b/Tests/feaLib/data/spec5f_ii_2.fea
@@ -3,7 +3,7 @@
 # http://www.adobe.com/devnet/opentype/afdko/topic_feature_file_syntax.html
 
 feature test {
-    @LETTER = [a-z];
+    @LETTER = [a - z];
     ignore sub @LETTER f' i';
     sub f' i' by f_i.begin;
 } test;
diff --git a/Tests/feaLib/data/spec5f_ii_3.fea b/Tests/feaLib/data/spec5f_ii_3.fea
index 5fd19916e..af06770de 100644
--- a/Tests/feaLib/data/spec5f_ii_3.fea
+++ b/Tests/feaLib/data/spec5f_ii_3.fea
@@ -3,7 +3,7 @@
 # http://www.adobe.com/devnet/opentype/afdko/topic_feature_file_syntax.html
 
 feature test {
-    @LETTER = [a-z];
+    @LETTER = [a - z];
     ignore sub @LETTER a' n' d', a' n' d' @LETTER;
     sub a' n' d' by a_n_d;
 } test;
diff --git a/Tests/feaLib/data/spec5f_ii_4.fea b/Tests/feaLib/data/spec5f_ii_4.fea
index 731a1f643..bc6fda43e 100644
--- a/Tests/feaLib/data/spec5f_ii_4.fea
+++ b/Tests/feaLib/data/spec5f_ii_4.fea
@@ -2,13 +2,13 @@
 # "Specifying exceptions to the Chain Sub rule"
 # http://www.adobe.com/devnet/opentype/afdko/topic_feature_file_syntax.html
 
-@LETTER = [A-Z a-z];
+@LETTER = [A - Z a - z];
 
 feature cswh {
 
     # --- Glyph classes used in this feature:
-    @BEGINNINGS = [A-N P-Z T_h m];
-    @BEGINNINGS_SWASH = [A.swash-N.swash P.swash-Z.swash T_h.swash m.begin];
+    @BEGINNINGS = [A - N P - Z T_h m];
+    @BEGINNINGS_SWASH = [A.swash - N.swash P.swash - Z.swash T_h.swash m.begin];
     @ENDINGS = [a e z];
     @ENDINGS_SWASH = [a.end e.end z.end];
 
diff --git a/Tests/feaLib/data/spec5fi3.fea b/Tests/feaLib/data/spec5fi3.fea
index e47a6f8aa..e44a7324b 100644
--- a/Tests/feaLib/data/spec5fi3.fea
+++ b/Tests/feaLib/data/spec5fi3.fea
@@ -5,5 +5,5 @@
 languagesystem latn dflt;
 
 feature test {
-    sub [A-Z] [A.sc-Z.sc]' by [a-z];
+    sub [A - Z] [A.sc - Z.sc]' by [a - z];
 } test;
diff --git a/Tests/feaLib/data/spec8a.fea b/Tests/feaLib/data/spec8a.fea
index b4d7dd299..405482137 100644
--- a/Tests/feaLib/data/spec8a.fea
+++ b/Tests/feaLib/data/spec8a.fea
@@ -10,7 +10,7 @@ feature aalt {
 } aalt;
 
 feature smcp {
-    sub [a-c] by [A.sc-C.sc];
+    sub [a - c] by [A.sc - C.sc];
     sub f i by f_i;  # not considered for aalt
 } smcp;